Meaning

• To step or walk (on or over something); to trample.

  "Don't tread on the lawn."

• To step or walk upon.

  "Actors tread the boards."

• To beat or press with the feet.

  "to tread a path; to tread land when too light; a well-trodden path"

• To go through or accomplish by walking, dancing, etc.

• To crush under the foot; to trample in contempt or hatred; to subdue.

• To copulate; said of (especially male) birds.

• (of a male bird) To copulate with.